Crossing-over contributes to the recombination of genetic material in offspring. When does crossing-over happen during meiosis?

Group of answer choices

when homologous chromosomes pair and portions of chromatids break off and are exchanged

when the DNA of the diploid cell is copied

when spindle fibers move the chromosomes toward the midline of the dividing cell

when homologous chromosomes move to opposite ends of the dividing cell