Maximum overlap is reached along with an active force in the skeletal muscle sarcomere. Due of titin's elastic recoil, passive increases. Actin/myosin overlap causes a decrease in active.
The distinct predictions of the tetanic force-length relationship of skeletal muscle made by the sliding filament and cross-bridge theories of muscle contraction have been put to the test in experiments. When the filament overlap is optimum, the active force produced by a maximally activated single fibre (with sarcomere length control) is optimal, and it decreases correspondingly when overlap is reduced. Because skeletal muscle has a static force-length relationship, it cannot foresee the effects of dynamic contractions. The active force is modulated during muscular contraction as a result of changes in sarcomere length, which is a phenomenon that is not always predicted by the cross-bridge theory.
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