There is some flexibility in pairing between the 3' base of the codon and the 5' base of the anticodons.
The typical genetic code uses fewer than 61 transfer RNAs to convert 61 codons into 20 amino acids (tRNAs). This is made feasible by the tRNA's capacity to "wobble" at the third nucleotide in order to decode several codons. Although the anticodon-codon mapping of tRNA to mRNA is a need for some codon usage indices and can help us understand the development of different genetic codes, such assays are frequently too expensive and complex to detect experimentally. Instead, using the wobble laws and theoretical deductions about nucleotide binding, the codon reading is approximated. Unfortunately, all of the subtleties of codon reading are not covered by these principles. This study examines the tRNAs' ability to read codons and how evolution has affected codon usage
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